Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)

Corresponding author: Francisco Brusquetti (franbrusquetti@gmail.com).

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1. autor: Brusquetti Estrada, Francisco Adolfo (author)
Kolejni autorzy: Bueno Villafañe, Diego (author), Caballero Gini, Andrea Natalia (author), Pereira Pinheiro, Paulo Durães (author), Netto Sisa, Flavia (author), Burgos Gallardo, Freddy (author), Fernández Ríos, Danilo (author), Cardozo Ruíz Díaz, Edgar Bernabé (author), Baldo, Diego (author)
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Wydane: 2025
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http://hdl.handle.net/20.500.14066/4682
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author Brusquetti Estrada, Francisco Adolfo
author2 Bueno Villafañe, Diego
Caballero Gini, Andrea Natalia
Pereira Pinheiro, Paulo Durães
Netto Sisa, Flavia
Burgos Gallardo, Freddy
Fernández Ríos, Danilo
Cardozo Ruíz Díaz, Edgar Bernabé
Baldo, Diego
author2_role author
author
author
author
author
author
author
author
author_browse Baldo, Diego
Brusquetti Estrada, Francisco Adolfo
Bueno Villafañe, Diego
Burgos Gallardo, Freddy
Caballero Gini, Andrea Natalia
Cardozo Ruíz Díaz, Edgar Bernabé
Fernández Ríos, Danilo
Netto Sisa, Flavia
Pereira Pinheiro, Paulo Durães
author_facet Brusquetti Estrada, Francisco Adolfo
Bueno Villafañe, Diego
Caballero Gini, Andrea Natalia
Pereira Pinheiro, Paulo Durães
Netto Sisa, Flavia
Burgos Gallardo, Freddy
Fernández Ríos, Danilo
Cardozo Ruíz Díaz, Edgar Bernabé
Baldo, Diego
author_role author
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dc.contributor.other.es.fl_str_mv Universidad Nacional de Asunción. Facultad de Ciencias Exactas y Naturales
dc.creator.none.fl_str_mv Brusquetti Estrada, Francisco Adolfo
Bueno Villafañe, Diego
Caballero Gini, Andrea Natalia
Pereira Pinheiro, Paulo Durães
Netto Sisa, Flavia
Burgos Gallardo, Freddy
Fernández Ríos, Danilo
Cardozo Ruíz Díaz, Edgar Bernabé
Baldo, Diego
dc.date.accessioned.none.fl_str_mv 2025-10-27T17:43:51Z
dc.date.available.none.fl_str_mv 2025-10-27T17:43:51Z
dc.date.issued.none.fl_str_mv 2025-10-17
dc.identifier.citation.en.fl_str_mv Brusquetti, F., Bueno-Villafañe, D., Caballero-Gini, A., Pinheiro, P. D. P., Netto, F., Burgos-Gallardo. F., Fernández Ríos, D., Cardozo, E. B., & Baldo, D. (2025) Unearthing the names of burrowing frogs: the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae). Vertebrate Zoology, 75, 325-352. https://doi.org/10.3897/vz.75.e159878
dc.identifier.doi.es.fl_str_mv 10.3897/vz.75.e159878
dc.identifier.essn.es.fl_str_mv 2625-8498
dc.identifier.issn.es.fl_str_mv 1864-5755
dc.identifier.other.es.fl_str_mv https://doi.org/10.3897/vz.75.e159878
dc.identifier.uri.none.fl_str_mv http://hdl.handle.net/20.500.14066/4682
dc.language.iso.es.fl_str_mv eng
dc.publisher.es.fl_str_mv Senckenberg Gesellschaft für Naturforschung
dc.relation.projectCONACYT.es.fl_str_mv PINV15-684
dc.rights.*.fl_str_mv Atribución/Reconocimiento 4.0 Internacional
dc.rights.accessRights.es.fl_str_mv info:eu-repo/semantics/openAccess
dc.rights.uri.*.fl_str_mv http://creativecommons.org/licenses/by/4.0/
dc.subject.classification.es.fl_str_mv 2. Medio ambiente
2.10. Protección de especies y sus hábitats
dc.subject.ocde.es.fl_str_mv 4. Ciencias Agrícolas y Veterinarias
4.1. Agricultura, silvicultura, pesca y ciencias afines (agronomía, zootecnia, pesca, silvicultura, horticultura, otras disciplinas afines)
dc.subject.other.es.fl_str_mv Dry Chaco
Endemic species
Humid Chaco
Leptodactylus gualambensis
dc.title.es.fl_str_mv Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)
dc.type.es.fl_str_mv info:eu-repo/semantics/article
info:eu-repo/semantics/publishedVersion
description Corresponding author: Francisco Brusquetti (franbrusquetti@gmail.com).
eu_rights_str_mv openAccess
format article
id CONACYT_0a246ce8da41d82adc6df9cde13ca929
identifier_str_mv Brusquetti, F., Bueno-Villafañe, D., Caballero-Gini, A., Pinheiro, P. D. P., Netto, F., Burgos-Gallardo. F., Fernández Ríos, D., Cardozo, E. B., & Baldo, D. (2025) Unearthing the names of burrowing frogs: the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae). Vertebrate Zoology, 75, 325-352. https://doi.org/10.3897/vz.75.e159878
1864-5755
10.3897/vz.75.e159878
2625-8498
language eng
network_acronym_str CONACYT
network_name_str Repositorio Institucional CONACYT
oai_identifier_str oai:repositorio.conacyt.gov.py:20.500.14066/4682
publishDate 2025
publishDateSort 2025
repository.mail.fl_str_mv repositorio.institucional@conacyt.gov.py
repository.name.fl_str_mv Repositorio Institucional CONACYT
repository_id_str
rights_invalid_str_mv Atribución/Reconocimiento 4.0 Internacional
http://creativecommons.org/licenses/by/4.0/
spelling 188600190600206600160f0155-e93e-4078-a5e1-6938ca640bfc600800600c09d7f8b-765f-474a-8bd5-f07a7e1294c9600419600de931612-fcba-4fad-81ef-6d633abda5216009a9ac526-cfeb-4ced-896f-671720615b1b600Universidad Nacional de Asunción. Facultad de Ciencias Exactas y Naturales2025-10-27T17:43:51Z2025-10-27T17:43:51Z2025-10-17Brusquetti, F., Bueno-Villafañe, D., Caballero-Gini, A., Pinheiro, P. D. P., Netto, F., Burgos-Gallardo. F., Fernández Ríos, D., Cardozo, E. B., & Baldo, D. (2025) Unearthing the names of burrowing frogs: the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae). Vertebrate Zoology, 75, 325-352. https://doi.org/10.3897/vz.75.e1598781864-5755https://doi.org/10.3897/vz.75.e159878http://hdl.handle.net/20.500.14066/468210.3897/vz.75.e1598782625-8498Corresponding author: Francisco Brusquetti (franbrusquetti@gmail.com).The widely distributed Neotropical frog Leptodactylus fuscus has long been suspected to represent a species complex. Among its numerous synonyms is Leptodactylus gualambensis Gallardo, 1964, described based on specimens from the western South American Gran Chaco and regarded as a typical Chacoan species. This species was synonymized 40 years ago and has not been reassessed since. In this study, we evaluate the taxonomic validity of the Chacoan populations of L. fuscus potentially assignable to L. gualambensis, based on molecular phylogenetic analyses, lineage delimitation, bioacoustics, and morphology using extensive sampling from the South American Gran Chaco and adjacent regions. Our phylogenetic analyses revealed three clades: One distributed across northern South America and Central America, including topotypes of L. fuscus; a second primarily occurring in the South American Gran Chaco, including topotypes of L. gualambensis; and a third found in parts of Paraguay, Argentina, and Brazil. This structure is corroborated by lineage delimitation analysis, genetic differentiation, gene flow estimations, and by differences in advertisement call duration and frequency, as well as head and limb proportions. Based on these results, we support the revalidation of L. gualambensis, redefine the geographic distribution of L. fuscus sensu stricto, and discuss the status of other available synonyms. In addition, we discuss variation in certain morphological traits of L. gualambensis that may be associated with its broad geographic distribution, which encompasses both dry, highly seasonal environments and humid, relatively homogeneous habitats.Consejo Nacional de Ciencia y TecnologíaPrograma Paraguayo para el Desarrollo de la Ciencia y Tecnología. Sistema Nacional de InvestigadoresPrograma Paraguayo para el Desarrollo de la Ciencia y Tecnología. Proyectos de investigación y desarrolloengSenckenberg Gesellschaft für NaturforschungAtribución/Reconocimiento 4.0 Internacionalhttp://creativecommons.org/licenses/by/4.0/info:eu-repo/semantics/openAccess© 2025 Francisco Brusquetti. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.2. Medio ambiente2.10. Protección de especies y sus hábitatsDry ChacoEndemic speciesHumid ChacoLeptodactylus gualambensis4. Ciencias Agrícolas y Veterinarias4.1. Agricultura, silvicultura, pesca y ciencias afines (agronomía, zootecnia, pesca, silvicultura, horticultura, otras disciplinas afines)Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)info:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersionVertebrate Zoology325352PINV15-68475Brusquetti Estrada, Francisco AdolfoBueno Villafañe, DiegoCaballero Gini, Andrea NataliaPereira Pinheiro, Paulo DurãesNetto Sisa, FlaviaBurgos Gallardo, FreddyFernández Ríos, DaniloCardozo Ruíz Díaz, Edgar BernabéBaldo, DiegoORIGINALUnearthing the names of burrowing frogs: the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae).pdfUnearthing the names of burrowing frogs: the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae).pdfArtículo científicoapplication/pdf7239682http://repositorio.conacyt.gov.py/bitstream/20.500.14066/4682/1/Unearthing%20the%20names%20of%20burrowing%20frogs%3a%20the%20taxonomic%20status%20of%20the%20Chacoan%20populations%20of%20Leptodactylus%20fuscus%20%28Schneider%2c%201799%29%20%28Anura%3a%20Leptodactylidae%29.pdfc3844354e13133a14e170bdfecdc650cMD51vertebrate_zoology-075-325-s001.pdfvertebrate_zoology-075-325-s001.pdfMaterial complementario 1application/pdf1387968http://repositorio.conacyt.gov.py/bitstream/20.500.14066/4682/4/vertebrate_zoology-075-325-s001.pdff6b48f9b150eaba28fc587c90cebf419MD54vertebrate_zoology-075-325-s002.pdfvertebrate_zoology-075-325-s002.pdfMaterial complementario 2application/pdf274678http://repositorio.conacyt.gov.py/bitstream/20.500.14066/4682/5/vertebrate_zoology-075-325-s002.pdfff99f26d7e3d2598131da6f28f1fffd7MD55vertebrate_zoology-075-325-s003.zipvertebrate_zoology-075-325-s003.zipMaterial complementario 3application/octet-stream75404http://repositorio.conacyt.gov.py/bitstream/20.500.14066/4682/6/vertebrate_zoology-075-325-s003.zip81603b044a5f7330a64f7905bc8d660bMD56CC-LICENSElicense_rdflicense_rdfapplication/rdf+xml; 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spellingShingle Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)
Brusquetti Estrada, Francisco Adolfo
2. Medio ambiente
2.10. Protección de especies y sus hábitats
Dry Chaco
Endemic species
Humid Chaco
Leptodactylus gualambensis
4. Ciencias Agrícolas y Veterinarias
4.1. Agricultura, silvicultura, pesca y ciencias afines (agronomía, zootecnia, pesca, silvicultura, horticultura, otras disciplinas afines)
status_str publishedVersion
title Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)
title_full Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)
title_fullStr Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)
title_full_unstemmed Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)
title_short Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)
title_sort Unearthing the names of burrowing frogs : the taxonomic status of the Chacoan populations of Leptodactylus fuscus (Schneider, 1799) (Anura: Leptodactylidae)
topic 2. Medio ambiente
2.10. Protección de especies y sus hábitats
Dry Chaco
Endemic species
Humid Chaco
Leptodactylus gualambensis
4. Ciencias Agrícolas y Veterinarias
4.1. Agricultura, silvicultura, pesca y ciencias afines (agronomía, zootecnia, pesca, silvicultura, horticultura, otras disciplinas afines)
url https://doi.org/10.3897/vz.75.e159878
http://hdl.handle.net/20.500.14066/4682